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I don't have an answer but never needed to know either. Only thing I can think of is it may have been needed for the mold or something since it provides no purpose I am aware of. If it was needed for headlight aiming, they would need two more notches for the High beams.
Like a guy scaring the **** out of me yesterday saying I got took on my NOS Bee grill as it has a notch out of the passenger side and not the drivers! I had to look...
Notch is so you can get the headlight retainer ring screw on that side..
Can't say on a 68, but on the 70s, there were provisions for grill mounting holes that the factory didn't use.
Putting my friends grill together, we decided to make them functional.